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Mapproxy-seed will seed each level independently for caches with upscale_tiles or downscale_tiles. You can use downscale_tiles to fill a cache for a source that only provides data for higher resolutions. It is therefore not recommended to use downscale_tiles values larger then one. A larger WMS request can quickly accumulate thousands of tiles required for downscaling. Downscaling from two levels below requires 16 tiles, three levels below requires 64, etc. Typically, a single tile can be downscaled from four tiles of the next zoom level. Note that the number of tiles growth exponentialy. The value configures by how many zoom levels MapProxy can search for a proper tile. MapProxy will scale down tiles from one or more zoom levels below (with higher resolutions) if you set downscale_tiles to 1 or higher. upscale_tiles allows you to provide responses for 10cm requests in all areas, allways returning the best available data. where you have an arial image cache of 20cm, with some areas also in 10cm resolution. This also works with partially seeded caches, eg. You can use upscale_tiles if you want to provide tiles or WMS responses in a higher resolution then your available cache. MapProxy will scale up tiles from one or more zoom levels above (with lower resolutions) if you set upscale_tiles to 1 or higher. MapProxy is able to create missing tiles by rescaling tiles from zoom levels below or above.